Permanent upgrades are purchased from the Ant Builder using Cornucopias. The cost of each upgrade starts at 3 Cornucopias and increases linearly at a rate of 1 up to a maximum of 20. Once the cost reaches 20, the ant will no longer offer additional upgrades. Some upgrades are unique and others are repeatable.
Some upgrades like "Water Pigeon" seem to be named, but most upgrades do not have a name currently.
Historian Ant
Keep a random upgrade from your previous run if you survived until fall.
Keep a random upgrade from your previous run if you survived past winter.
Start with 1 additional seed for each season survived last run.
Start next run with an random farm animal if you survived until winter in your previous run.
Gain 2 heart sprouts at the start of your next run if you survived until Summer in your previous run.
Start with an additional 2 fertilizer for each season you survived last run.
Start with an additional pickaxe.
Increase Sprinkler Tractor duration and watering radius